Schedule, syllabus and examination date

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Course content

The course provides a thorough introduction to modern evolutionary science, with an emphasis on phenotypic evolution, life history theory, adaptation, speciation and macroevolution.

Learning outcome

After completing the course you have:

  • Good understanding of basic concepts within the theory of evolution such as phylogeny, natural selection, fitness, plasticity, adaptation, speciation, inheritance and genetic variation.
  • Good understanding of the main principles of quantitative genetics so that you can perform calculations of the evolution of phenotypic characters.
  • Knowledge of how different life history strategies (organism growth, development and reproduction) can be explained through adaptation and natural selection.
  • Knowledge of the processes that give rise to new species, the main features of the evolution of life on Earth and the factors that determine the geographic distribution of organisms.
  • Insight into how evolution takes place over different time scales, and how micro- and macroevolution is linked.
  • Gained experience in finding, acquainting yourself with, critically evaluating and disseminating research literature in writing and orally.


Students who are admitted to study programmes at UiO must each semester register which courses and exams they wish to sign up for in Studentweb.

If you are not already enrolled as a student at UiO, please see our information about admission requirements and procedures.

Priority will be given to students in the bachelor program in Biosciences.


Formal prerequisite knowledge

In addition to fulfilling the Higher Education Entrance Qualification, applicants have to meet the following special admission requirements:

  • Mathematics R1 (or Mathematics S1 and S2) + R2

And in addition one of these:

  • Physics (1+2)
  • Chemistry (1+2)
  • Biology (1+2)
  • Information technology (1+2)
  • Geosciences (1+2)
  • Technology and theories of research (1+2)

The special admission requirements may also be covered by equivalent studies from Norwegian upper secondary school or by other equivalent studies (in Norwegian).

Recommended previous knowledge

Overlapping courses


  • Lectures
  • Qolloquia/computer labs/seminars with calculations
  • Research paper presentation (obligatory)
  • Essay Writing 6 pages (obligatory)

Attendance to the first lecture is mandatory, also for those on the waiting list. You will lose your seat on the course if documentation for valid absence is not provided to the student administration prior to the first lecture.


  • Mandatory essay counts for 30% of the mark
  • Final written exam counts for 70% of the mark

The research paper presentation must be approved before the exam

Examination support material

No examination support material is allowed.

Language of examination

The examination text is given in English.You may submit your response in Norwegian, Swedish, Danish or English.

Grading scale

Grades are awarded on a scale from A to F, where A is the best grade and F is a fail. Read more about the grading system.

Explanations and appeals

Resit an examination

This course offers both postponed and resit of examination. Read more:

Withdrawal from an examination

It is possible to take the exam up to 3 times. If you withdraw from the exam after the deadline or during the exam, this will be counted as an examination attempt.

It will also be counted as one of the three attempts to sit the exam for this course, if you sit the exam for one of the following courses:  BIOS4100 – Evolution

Special examination arrangements

Application form, deadline and requirements for special examination arrangements.


The course is subject to continuous evaluation. At regular intervals we also ask students to participate in a more comprehensive evaluation.

Facts about this course






Every spring

The course is offered for the first time in 2019


Every spring

Teaching language

Norwegian (English on request)